Perennials 15-100+ cm (caudices or taproots woody). Stems erect, branched from bases. Leaves basal; alternate; petiolate or sessile; blades 3-nerved, elliptic, ovate, rhombic, or suborbiculate, bases ± cuneate to nearly truncate, margins entire (sometimes corrugate or ruffled), faces densely puberulent or silky-velutinous. Heads radiate, borne singly (peduncles much longer than involucres). Involucres ± hemispheric or broader, 10-30+ mm diam. Phyllaries persistent, 30-65+ in 3-6 series. Receptacles convex, paleate (paleae ± conduplicate, folded around and falling with cypselae). Ray florets (11-)20-35+, neuter; corollas yellow. Disc florets (50-)200-500+, bisexual, fertile; corollas yellow, tubes shorter than to equaling abruptly expanded throats, lobes 5, triangular. Cypselae strongly compressed or flattened, ± cuneate (margins ± white, corky, usually ciliate, faces black, glabrous or ± silky-villous); pappi 0, or persistent, of 2 awns (often with 2-10+, often connate, minute scales or teeth as well). x = 18.