From protologue (Sida. 21 (1): 210. 2004): Keil 28401 (holotype- flowering stem of plant D, OBI; ISOTYPES [the letter in parentheses represents the individual], ASU (A), BRY (C), CAS (B), CDA (C), DAO (D), GH (D), MIN (D), MO (C), NY (C). OBI (A, C), RSA (A), TEX (D), UC (B), US (D))